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MacLeod is picked up at the airport in France by his old friend, Hugh Fitzcairn. For the first time in the 350 years MacLeod has known him. Fitz has settled down as a Universitv Professor with the love of his life, Naomi. Flashbacks show MacLeod and Fitz's first meeting, when MacLeod was protecting the Doge's daughter from Fitz's amourous advances in Venice, 1637. When Naomi's jealous ex-lover is found strangled next to a computer displaying Fitz's falsified teaching credentials, Fitz goes on the run from the police. MacLeod realizes that Fitz is being framed by Kalas, who has followed him from the U.S. Kalas challenges Fitz and takes his head while MacLeod watches, unable to interfere.

In the 1600s, MacLeod sought refuge in a monastery founded by Paul, another Immortal. There he encountered Kalas, an Immortal monk with a heavenly singing voice. When it was discovered that Kalas was routinely taking the heads of Immortals as they left the sanctuary, he was expelled from the monastery and the music that was his life. Now, Paul and his choir have been lured out of their monastery for a concert tour and MacLeod discovers that Kalas is after revenge. Meanwhile, mysterious deaths at the hospital seem linked to negligence on Anne's part. It becomes obvious that Kalas is trying to destroy MacLeod's friends before coming for him.

When a religious leader, John Kirin, dies on Anne's operating table and then returns from the dead, his believers know a miracle has occurred. MacLeod knows better. He watched as Kirin, then known as Kage, massacred POWs in the Spanish Civil War and left a band of Cambodian refugee children to die at the hands of the Khmer Rouge. Kirin swears that experience changed him forever, turning him from a man of war to a man of peace. When a tabloid reporter trying to get the goods on Kirin winds up dead in MacLeod's dojo, MacLeod is certain Kirin is responsible. Kirin protests his innocence and realizes the real killer is Matthew, one of his faithful disciples trying to protect him. Kirin confronts a disillusioned Matthew, who manages to kill Kirin before dying himself in a rain of police bullets. In the Tag, Kirin and MacLeod have made peace as Kirin takes to the road, hoping to do good elsewhere.

Recent Immortal Michael Christian has been on an incredible string of luck, taking a number of heads from unarmed and vulnerable opponents, including May-Ling Shen, who taught MacLeod the martial arts in 1780's Mongolia. Christian's Watcher, Rita Luce, has been doing more than just watching, supplying Christian with classified information on the other Immortals and their weaknesses. MacLeod, unaware of Christian, goes on a vision quest to his cabin on Holy Ground — deliberately leaving his sword behind. The race is on for Joe Dawson to figure out Rita's secret before Christian sets his sights on MacLeod.

To save his own hide, petty hoodlum Benny Carbassa, the Nathan Detroit of Immortality, turns MacLeod over to an aging gangster determined to see MacLeod dead before he dies. In the midst of this, Anne returns, having convinced herself that MacLeod will open up in his own time and determined not to push him too hard. In flashback, we see MacLeod's first meeting with Benny, in 1938 at the Coconut Lounge, a club operated by two young brothers who are rivals for the same torch singer.