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MacLeod is struck and "killed" by a speeding automobile. When he revives in the hospital, he attracts the attention of a deranged doctor, who kidnaps MacLeod for his research on a miracle cure for trauma. When MacLeod escapes from the doctor, the doctor murders a potential witness and frames MacLeod for it.

When Tessa is taken by a band of Survivalists who live off the land in the vastness of the Pacific Northwest, MacLeod must call on all of his own skills, honed a hundred and fifty years earlier, when he roamed this same mountain vastness. The leader of these modern day mountain men, Caleb (who is revealed to be an Immortal), falls in love with Tessa, and in the end he and MacLeod battle to the death, axe against sword, against the primitive backdrop of the timeless wilderness.

While visiting the courthouse to clear up Tessa's traffic tickets, MacLeod, Tessa, and Richie are among a group that is taken hostage by a band of men bent on freeing their leader, Slade, from imprisonment. When Slade's demands are not met, he chooses a hostage to sacrifice. MacLeod makes sure that hostage is himself, and returns from the dead to hunt down the gunmen one by one, as the police watch helplessly from outside.

A beautiful young woman, Felicia Martins, throws herself to her death, only to revive with the realization that she is an Immortal. She seeks out Duncan MacLeod as her mentor, and becomes amorously involved with Richie Ryan. But there is much more to this "ingénue" than meets the eye. She is in fact a long-time Immortal, engaged in a wily scheme to trap MacLeod and acquire his Quickening. MacLeod must see through her subterfuge in time to protect not only himself, but also Richie.

An Immortal friend of MacLeod's is beheaded in his isolated mountain retreat where he had hoped – like MacLeod – to retire from the fight for a while. A brain-damaged, homeless veteran is charged with the grisly crime, and an attempt is made to link him to the killing of Slan Quince (the villain of episode one, 'The Gathering'), an Evil Immortal whom MacLeod himself had slain in combat. MacLeod must try and clear the innocent man and unravel the mystery of who killed his friend. Here again we get a glimpse of MacLeod's past, in flashbacks set during the American Civil War.