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Donna, a girl Richie dated briefly several years ago, returns with her 18-month-old son — who she claims is Richie's! While MacLeod reminds him that it's impossible, that Immortals are unable to have children, Richie sees this as an opportunity to have the family he never had and never will be able to again. When Kern, an evil Immortal, rides into town, MacLeod is reminded of his own foster son, Kahani. Kahani and his Sioux Indian mother, Little Deer, were massacred over a hundred years ago by U.S. soldiers led by Kern, then a mercenary scout. MacLeod is eager to even the score. When Donna finds Richie's sword and demands to know what it's for, Richie is at a loss to tell her. MacLeod advises Richie that it would be safer and kinder to Donna and the baby to leave them. When his new-found family is threatened by Kern, Richie realizes MacLeod is right. While MacLeod rids the world of the evil Kern, Richie lets go of the only family he'll ever have.

After Midori Koto sees her husband, rich industrialist Michael Kent, murder her lover, she kills Kent and runs to MacLeod for protection. She reminds MacLeod of a vow of protection his "ancestor" (MacLeod himself) made 200 years before. Flashbacks tell the story of MacLeod coming to the aid of the samurai Hideo Koto after MacLeod is shipwrecked in Japan. He vows to Hideo he will always protect the Koto family and is bequeathed the dragon head katana sword he uses to this day. Back in the present, MacLeod discovers that Kent is an Immortal and he's still alive.

Joe Dawson arrives in Paris to let MacLeod know that Horton may still be alive and coming after him. However, when MacLeod meets ""Lisa Milon,"" a dead-ringer for Tessa (Lisa Halle after plastic surgery), all thoughts of Horton and the Watchers go out of his head. Richie, unsure if Mac is thinking clearly, goes to Dawson for help and winds up taking a bullet for him when Horton tries a hit. Lisa is kidnapped, and MacLeod insists on going after her even though his friends warn him it may be a setup. He can't take a chance on burying her again. When Lisa finally tries to kill MacLeod, he turns the tables on her and Horton at long last gets what's coming to him.

Pete Wilder just saved Richie from an attack by some guys wearing Watcher tattoos. Or did he? Nothing is what it seems, and MacLeod becomes increasingly wary of Pete, despite Richie's protests that the guy is his friend. When MacLeod's mistrust gets the possibly innocent Pete killed, Mac becomes increasingly unsure of himself. Unknown to MacLeod, his old enemy Horton is pulling all the strings. Horton is secretly training escaped killer Lisa Halle to go after MacLeod while he's troubled and vulnerable.