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Father Bernard has a secret he thought he'd buried 50 years ago at the bottom of the Seine. When Ernest Daimler, the Nazi Major who Bernard killed as a young boy and threw in the river, appears at his church looking not a day older than the day he died, Father Bernard realizes he's like MacLeod. As a child, Bernard watched MacLeod die and revive during a mission for the French Resistance and MacLeod swore him to secrecy. Bernard goes to MacLeod for help. Meanwhile, Anne tells MacLeod that she is pregnant — by an old friend she sought for comfort after MacLeod's ""death."" MacLeod tries to adjust to the concept of being a father. When Daimler kills Father Bernard and comes after Anne, MacLeod kills Daimler and Anne witnesses what being part of MacLeod's life really means. Unable to deal with the part of herself that wanted to see Daimler die, Anne leaves MacLeod.

MacLeod decides to tell Anne the truth about his Immortality and she flies to Paris to be with him. En route, Anne helps save the life of a young woman, Tasha, who turns out to be smuggling drugs for the Russian Mafia. Tasha is the lover of Kristov, the head of the Russian gang and formerly the leader of the band of Cossacks MacLeod encountered on his way to the Orient in 1750. Concerned about Tasha, Anne tries to convince her to testify against Kristov, while Kristov is determined to make sure Tasha dies before she can testify. Richie is kidnapped by Kristov as a pawn in this game. MacLeod must choose between taking down Kristov or saving Richie and Richie finds he must grow as an Immortal — or die.

When Immortal Ceirdwyn and her mortal husband are gunned down by a street gang, she calls upon her skills as an ancient Celtic warrior to exact her revenge on the members of the gang, one by one. At the racetrack watching Richie's success at racing, MacLeod befriends a young pickpocket, the brother of one of the gang members, and learns of the killings. MacLeod, who has known Ceirdwyn since before they helped smuggle Bonnie Prince Charlie out of Scotland in 1746, feels he must stop Ceirdwyn and make her see that revenge is not the answer — a lesson she helped MacLeod learn in the bloody aftermath of Culloden. In return, Ceirdwyn helps MacLeod see that, although loving a mortal can be dangerous for the mortal, it is the mortal who must choose whether to take the risk. MacLeod calls Anne. Meanwhile, Richie ""dies"" in a firey crash during a race, a crash that also takes the life of the champion, Basil.